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mailflow broken - error 451 4.7.0

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Hi,

i have the following situation with an exchange 2010/2013 coexistence scenario:

3 Servers, 1 2008 R2 Server as DC with Exchange 2010, 1 2012 Server as DC, 1 2012 Server with Exchange 2013. The Problem is that the mailflow on the 2013 Exchange is simply not working, and I have tried all suggested solutions i could find in the last 2 days without success. 
Test-Mailflow fails while Test-SmtpConnectivity succeeds, but 2013-mailboxes cannot receive any mail. SmtpReceive Logs are like this:

2013-04-13T16:41:12.526Z,EX01\Default Frontend EX01,08D004497C6E9CC3,25,127.0.0.1:25,127.0.0.1:61348,*,,Message or connection acked with status Retry and response 451 4.4.0 DNS query failed. The error was: DNS query failed with error ErrorRetry
2013-04-13T16:41:12.526Z,EX01\Default Frontend EX01,08D004497C6E9CC3,26,127.0.0.1:25,127.0.0.1:61348,>,451 4.7.0 Temporary server error. Please try again later. PRX2 ,

i get the same errors when i telnet from the 2010 to the 2013 server.
so it seems like a DNS related problem, but i cannot figure out whats the problem. the ex2013 has the 2 DCs as internal DNS servers and i tried all 3 possible options of the DNS lookups setting in the server properties in ECP. i can resolve all servers without problems on any server, so i really dont understand whats the problem here...

any ideas? somebody? please  ;)

edit:

heres another snippet from the connectivity-log:

2013-04-13T23:59:44.720Z,08D006869E9DDB49,SMTP,internalproxy,>,DNS server returned ErrorRetry reported by 0.0.0.0. [Domain:Result] = EX01.xyz.priv:ErrorRetry;
2013-04-13T23:59:44.720Z,08D006869E9DDB49,SMTP,internalproxy,-,Messages: 0 Bytes: 0 (The DNS query for  'Undefined':'internalproxy':'00000000-0000-0000-0000-000000000000' failed with error : ErrorRetry)
2013-04-14T00:00:46.316Z,08D006869E9DDB4C,SMTP,internalproxy,+,Undefined 00000000-0000-0000-0000-000000000000;QueueLength=0
2013-04-14T00:01:46.404Z,08D006869E9DDB4C,SMTP,internalproxy,>,DNS server returned ErrorRetry reported by 0.0.0.0. [Domain:Result] = EX01.xyz.priv:ErrorRetry;
2013-04-14T00:01:46.404Z,08D006869E9DDB4C,SMTP,internalproxy,-,Messages: 0 Bytes: 0 (The DNS query for  'Undefined':'internalproxy':'00000000-0000-0000-0000-000000000000' failed with error : ErrorRetry)

so the dns queries for 'undefined' and 'internalproxy' are failing, but i really don't know what that means.



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